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Publications and Educational material by the Cephalopod Citizen Science Project available for download.


Drerup, C. and Cooke, G.M., 2021. Shoaling behaviour in the European cuttlefish Sepia officinalis. Ethology, 127(12), pp.1101-1108.

Drerup, C., Jackson, A., Rickard, C., Skea, M. and Cooke, G.M., 2021. Field observations on the behavioural ecology of the stout bobtail squid Rossia macrosoma (Cephalopoda: Sepiolidae) from Scottish waters.​

Laptikhovsky, V., Cooke, G., Barrett, C., Lozach, S., MacLeod, E., Oesterwind, D., Sheerin, E., Petroni, M., Barnwall, L., Robin, J.P. and Allcock, L., 2021. Identification of benthic egg masses and spawning grounds in commercial squid in the English Channel and Celtic Sea: Loligo vulgaris vs L. forbesii. Fisheries Research, 241, p.106004.

Gordon, D., Pugh, P. and Cooke, G.M., 2019. Social Media and citizen science provide valuable data for behavioural ecology research: Are cuttlefish using pursuit-deterrent signals during hunting?. bioRxiv, p.760926.


Oliveira, A.R., Sardinha-Silva, A., Andrews, P.L., Green, D., Cooke, G.M., Hall, S., Blackburn, K. and Sykes, A.V., 2020. Microplastics presence in cultured and wild-caught cuttlefish, Sepia officinalis. Marine Pollution Bulletin, 160, p.111553.

Sykes, A.V., Almansa, E., Ponte, G., Cooke, G.M. and Andrews, P.L., 2020. Can cephalopods vomit? Hypothesis based on a review of circumstantial evidence and preliminary experimental observations. Frontiers in Physiology, 11.

Drerup, C., Sykes, A.V. and Cooke, G.M., 2020. Behavioural aspects of the spotty bobtail squid Euprymna parva (Cephalopoda: Sepiolidae). Journal of Experimental Marine Biology and Ecology, 530, p.151442.

Cooke, G.M., Anderson, D.B., Begout, M.L., Dennison, N., Osorio, D., Tonkins, B., Kristiansen, T., Fiorito, G., Galligioni, V., Ponte, G. and Andrews, P.L., 2019. Prospective severity classification of scientific procedures in cephalopods: Report of a COST FA1301 Working Group survey. Laboratory animals, p.0023677219864626.

Cooke, G.M., Tonkins, B.M. and Mather, J.A., 2019. Care and Enrichment for Captive Cephalopods. In The Welfare of Invertebrate Animals (pp. 179-208). Springer, Cham.


Sykes, António V., Eduardo Almansa, Gavan M. Cooke, Giovanna Ponte, and Paul LR Andrews. “The digestive tract of cephalopods: a neglected topic of relevance to animal welfare in the laboratory and aquaculture.” Frontiers in Physiology 8 (2017): 492.

Cooke, G. M & Tonkins, B. M. 2015 Behavioural indicators of welfare exhibited by the common European cuttlefish (Sepia officinalis). Journal of Zoo and Aquarium Research 3(4)

Tonkins, Belinda M., Alexandra M. Tyers, and Gavan M. Cooke. 2015 “Cuttlefish in captivity: An investigation into housing and husbandry for improving welfare.” Applied Animal Behaviour Science 168 (77-83).


National Geographic SEE Initiative ROV Drone grant win. Value ~$1600 The secret Lives of shell dwelling octopuses

National Geographic SEE Initiative ROV Drone grant win. Value ~$1600 How to film cephalopods


CIAC/Cephs in Action conference, Heraklion, Crete March 2017 Presenting talk: “Prospective severity assessment in cephalopods: Results of an on-line survey of the COST Action (CephsInAction) FA1301 cephalopod research community”

Aquaculture Europe, 2016, Edinburgh, oral presentation: “Effects of increased volume and tank bottom areas on cuttlefish Sepia officinalis, reproduction behaviour”

RSPCA/APHA ‘Welfare of wild animals used in research’ meeting, 2016, oral presentation: “Recent Inclusion of the Class Cephalopoda (Mollusca) in ASPA1986/EU Directive 2010/63/EU and The Use of Wild Caught Specimens in Scientific Procedures”

LASA/UFAW meeting, 2016, oral presentation: Welfare assessment in alien life forms: The challenges presented by inclusion of cephalopods in directive 2010/63/EU

BBSRC Animal Welfare Research Network Inaugural Meeting, May 2016, Welcome Trust, London, poster: “COST Action FA1301 – CephsInAction A network for improvement of cephalopod welfare and husbandry in research, aquaculture and fisheries”

FELASA congress 2016, Brussels, poster: “Prospective severity assessment in cephalopods: Results of an on-line survey of the COST Action (CephsInAction) FA1301 cephalopod research community”
Aquaculture Europe, 2015, Rotterdam, oral presentation: “Caring for cephalopods”